1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;3 Hebei Academy of Forestry,Shijiazhuang 050061,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;,1Institute of Forest Biotechnology, Forestry College, Agricultural University of Hebei, Baoding 071000 ;2 Hebei Key Laboratory for Tree Genetic Resources and Forest Protection, Baoding 071000;